Complex II does not pump protons across the mitochondrial inner membrane when it is actively transferring electrons from its substrate to its product.
Complex II is unique. Unlike other Complexes- I, III, and IV, complex II takes part in the electron transport chain but it does not actively pump protons across the inner mitochondrial membrane. With each electron transport, the electron transport chain releases minute amounts of energy and this transport is coupled to the pumping of protons across the mitochondrial membrane. Eventually, this electrons are delivered to molecular oxygen. Which is then reduced to water, in Complex IV. Complex II is the enzyme that carries out this transformation in the citric acid cycle. That is why the complex II is also known as succinate dehydrogenase.
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