In this excerpt from the poem "Old Ironsides" by Oliver Wendell Holmes, what is the meaning of the word tread?

Her deck, once red with heroes' blood
Where knelt the vanquished foe,
When winds were hurrying o'er the flood
And waves were white below,
No more shall feel the victor's tread,
Or know the conquered knee;—

A.
crush
B.
pace
C.
quell
D.
trudge
E.
march